Team newsThe Ireland coach has restored Jamison Gibson-Park, Tadhg Furlong, Tadhg Beirne and Josh van der Flier to the team that beat Italy, with Cormac Izuchukwu and Edwin Edogbo missing out on the matchday 23.
Crowley impressed off the bench last week and is backed for his first start since the defeat to New Zealand in Chicago in November.
Henry Pollock will make his first England start in Saturday’s Guinness Six Nations clash with Ireland as one of three changes in personnel to the side defeated by Scotland.
Pollock lines up at number eight as part of a reshaped back row that sees Ben Earl move to openside and Tom Curry come in at blindside, with Sam Underhill and Guy Pepper dropping to the bench.
England
Freddie Steward, Tommy Freeman, Ollie Lawrence, Fraser Dingwall, Henry Arundell, George Ford, Alex Mitchell, Ellis Genge, Luke Cowan-Dickie, Joe Heyes, Maro Itoje, Ollie Chessum, Tom Curry, Ben Earl, Henry Pollock
Replacements: Jamie George, Bevan Rodd, Trevor Davison, Alex Coles, Guy Pepper, Sam Underhill, Jack van Poortvliet, Marcus Smith
Ireland: Jamie Osborne; Rob Baloucoune, Garry Ringrose, Stuart McCloskey, James Lowe; Jack Crowley, Jamison Gibson-Park; Jeremy Loughman, Dan Sheehan, Tadhg Furlong, James Ryan, Joe McCarthy, Tadhg Beirne, Josh van der Flier, Caelan Doris (capt).
Replacements: Rónan Kelleher, Tom O’Toole, Finlay Bealham, Jack Conan, Nick Timoney, Craig Casey, Ciarán Frawley, Tommy O’Brien.