Lima LJR, Almeida MH, Nout MJR, Zwietering MH. Theobroma cacao L., the food of the gods: quality determinants of commercial cocoa beans, with particular reference to the impact of fermentation. Crit Rev Food Sci Nutr. 2011;51:731–61.
Niemenak N, Cilas C, Rohsius C, Bleiholder H, Meier U, Lieberei R. Phenological growth stages of cacao plants (Theobroma sp.): codification and description according to the BBCH scale. Ann Appl Biol. 2010;156:13–24.
Beg MS, Ahmad S, Jan K, Bashir K. Status, supply chain and processing of cocoa – a review. Trends Food Sci Technol. 2017;66:108–16.
Baruah IK, Shao J, Ali SS, Schmidt ME, Meinhardt LW, Bailey BA, et al. Cacao pod transcriptome profiling of seven genotypes identifies features associated with post-penetration resistance to Phytophthora palmivora. Sci Rep. 2024;14:4175.
Chepsergon J, Moleleki LN. Rhizosphere bacterial interactions and impact on plant health. Curr Opin Microbiol. 2023;73: 102297.
Ploetz RC. Cacao diseases: important threats to chocolate production world wide. Phytopathology. 2007;97:1634–9.
Madhu GS, Rani AT, Muralidhara BM, Rajendiran S, Venkataravanappa V, Sriram S. First report of Phytophthora tropicalis causing black pod of cacao (Theobroma cacao) in India. Australas Plant Pathol. 2023;52:591–3.
Acebo-Guerrero Y, Hernández-RodrÃguez A, Heydrich-Pérez M, El Jaziri M, Hernández-Lauzardo AN. Management of black pod rot in cacao (Theobroma Cacao L.): a review. Fruits. 2012;67:41–8.
Doungous O, Minyaka E, Longue EAM, Nkengafac NJ. Potentials of cocoa pod husk-based compost on phytophthora pod rot disease suppression, soil fertility, and Theobroma Cacao L. growth. Environ Sci Pollut Res. 2018;25:25327–35.
Miguelez-Sierra Y, Acebo-Guerrero Y, El Jaziri M, Bertin P, Hernández-RodrÃguez A. Pseudomonas chlororaphis CP07 strain reduces disease severity caused by Phytophthora palmivora in genotypes of Theobroma cacao. Eur J Plant Pathol. 2019;155:1133–43.
Gupta A, Gopal M, Thomas GV, Manikandan V, Gajewski J, Thomas G, et al. Whole genome sequencing and analysis of plant growth promoting bacteria isolated from the rhizosphere of plantation crops coconut, cocoa and arecanut. PLoS One. 2014;9:e104259.
Beneduzi A, Ambrosini A, Passaglia LMP. Plant growth-promoting rhizobacteria (PGPR): their potential as antagonists and biocontrol agents. Genet Mol Biol. 2012;35(4 suppl 1):1044–51.
Hasan A, Tabassum B, Hashim M, Khan N. Role of plant growth promoting rhizobacteria (PGPR) as a plant growth enhancer for sustainable agriculture :a Review. Bacteria. 2024:3(2):59–75. https://doi.org/10.3390/bacteria3020005.
Nagargade M, Tyagi V, Singh MK. Plant Growth-Promoting rhizobacteria: A biological approach toward the production of sustainable agriculture. In: Meena VS, editor. Role of rhizospheric microbes in soil. Singapore: Springer Singapore; 2018. pp. 205–23.
Reddy PP. Potential role of PGPR in agriculture. Plant growth promoting rhizobacteria for horticultural crop protection. New Delhi: Springer India; 2014. pp. 17–34.
Aarti S, Parmar S. Plant growth promoting rhizobacteria: their potential in sustainable agriculture. J Sci Innov Nat Earth. 2024;4:40–6.
Lugtenberg B, Kamilova F. Plant-growth-promoting rhizobacteria. Annu Rev Microbiol. 2009;63:541–56.
Widyanta Pratama S, Sukamto S, Nur Asyiah L, Vida Ervina Y. Growth Inhibition of cocoa pod rot fungus phytophthora palmivora by Pseudomonas fluorescence and Bacillus subtilis bacteria. Pelita Perkeb. 2013:29 (2):120–127. https://doi.org/10.22302/iccri.jur.pelitaperkebunan.v29i2.59.
Jibat M, Alo S. Integrated management of black pod (Phytophthora palmivora) disease of cocoa through fungicides and cultural practices in Southwestern Ethiopia. IJ-FANRES. 2023;4:43–5.
McGREGOR AJ. A small-scale screening technique for evaluating fungicides against Phytophthora palmivora pod rot of cocoa. Ann Appl Biol. 1982;101:25–31.
Sowunmi FA, Famuyiwa GT, Oluyole KA, Aroyeun SO, Obasoro OA. Environmental burden of fungicide application among cocoa farmers in Ondo state, Nigeria. Sci Afr. 2019;6: e00207.
De Souza JT, De Boer M, De Waard P, Van Beek TA, Raaijmakers JM. Biochemical, genetic, and zoosporicidal properties of cyclic lipopeptide surfactants produced by Pseudomonas fluorescens. Appl Environ Microbiol. 2003;69:7161–72.
Harman GE, Howell CR, Viterbo A, Chet I, Lorito M. Trichoderma species — opportunistic, avirulent plant symbionts. Nat Rev Microbiol. 2004;2:43–56.
Choudhary DK, Johri BN. Interactions of Bacillus spp. and plants – with special reference to induced systemic resistance (ISR). Microbiol Res. 2009;164:493–513.
Madhu GS, Rani AT, Muralidharan BM, Rajendiran S. A modified fungicide based media for the isolation of Phytophthora Cinnomomi rands. Causing avocado root rot. Pest Manage Hortic Ecosyst. 2023;23:302–3.
Abad ZG, Burgess TI, Redford AJ, Bienapfl JC, Srivastava S, Mathew R, et al. IDphy: an international online resource for molecular and morphological identification of Phytophthora. Plant Dis. 2023;107:987–98.
Delgadillo-Durán P, Soto-Suárez M, Rodriguez-Polanco L, Carrero-Gutierrez M, Torres-Rojas E, Yockteng R. A new method for the inoculation of Phytophthora palmivora (Butler) into cacao seedlings under greenhouse conditions. Plant Methods. 2020;16: 114.
Tamura K, Stecher G, Kumar S. MEGA11: molecular evolutionary genetics analysis version 11. Mol Biol Evol. 2021;38:3022–7.
Bergey DH, Holt JG, Krieg NR, editors. Bergey’s manual of determinative bacteriology. 9th ed. Baltimore: Williams & Wilkins; 1994.
Taylor WI, Achanzar D. Catalase test as an aid to the identification of Enterobacteriaceae. Appl Microbiol. 1972;24:58–61.
Jurtshuk P, McQuitty DN. Use of a quantitative oxidase test for characterizing oxidative metabolism in bacteria. Appl Environ Microbiol. 1976;31:668–79.
Kanlaya R, Subkod C, Thongboonkerd V. A novel, simple and rapid assay to measure citrate level in bacterial culture for analysis of citrate consumption by bacteria. Talanta Open. 2024;10:100360.
Patten CL, Glick BR. Role of Pseudomonas putida indoleacetic acid in development of the host plant root system. Appl Environ Microbiol. 2002;68:3795–801.
Kifle MH, Laing MD. Isolation and screening of bacteria for their diazotrophic potential and their influence on growth promotion of maize seedlings in greenhouses. Front Plant Sci. 2016;6:1225. https://doi.org/10.3389/fpls.2015.01225.
Schwyn B, Neilands JB. Universal chemical assay for the detection and determination of siderophores. Anal Biochem. 1987;160:47–56.
Murphy J, Riley JP. A modified single solution method for the determination of phosphate in natural waters. Anal Chim Acta. 1962;27:31–6.
O’Toole GA. Microtiter dish biofilm formation assay. J Vis Exp. 2011;30(47):2437. https://doi.org/10.3791/2437.
Wheeler BEJ. An introduction to plant diseases. Wiley and Sons, London. 1969;374.
Wilcoxson RD, Skovmand B, Atif AH. Evaluation of wheat cultivars for ability to retard development of stem rust. Ann Appl Biol. 1975;80:275–81.
Glick BR, Todorovic B, Czarny J, Cheng Z, Duan J, McConkey B. Promotion of plant growth by bacterial ACC deaminase. Crit Rev Plant Sci. 2007;26:227–42.
Gerhardt KE, Huang X-D, Glick BR, Greenberg BM. Phytoremediation and rhizoremediation of organic soil contaminants: potential and challenges. Plant Sci. 2009;176:20–30.
Bhattacharyya PN, Jha DK. Plant growth-promoting rhizobacteria (PGPR): emergence in agriculture. World J Microbiol Biotechnol. 2012;28:1327–50.
Vacheron J, Desbrosses G, Bouffaud M-L, Touraine B, Moënne-Loccoz Y, Muller D, et al. Plant growth-promoting rhizobacteria and root system functioning. Front Plant Sci. 2013. https://doi.org/10.3389/fpls.2013.00356.
Nepali B, Bhattarai S, Jiban S. Identification of Pseudomonas fluorescens using different biochemical tests. 2020. https://doi.org/10.13140/RG.2.2.23860.40328
Lalucat J, Bennasar A, Bosch R, GarcÃa-Valdés E, Palleroni NJ. Biology of Pseudomonas stutzeri. Microbiol Mol Biol Rev. 2006;70:510–47.
Hallmann J, Berg G. Spectrum and population dynamics of bacterial root endophytes. In: Schulz BJE, Boyle CJC, Sieber TN, editors. Microbial root endophytes. Berlin, Heidelberg: Springer Berlin Heidelberg; 2006. pp. 15–31.
Zahid M, Abbasi MK, Hameed S, Rahim N. Isolation and identification of Indigenous plant growth promoting rhizobacteria from Himalayan region of Kashmir and their effect on improving growth and nutrient contents of maize (Zea Mays L). Front Microbiol. 2015;6:207. https://doi.org/10.3389/fmicb.2015.00207.
Hardiansyah MY, Musa Y, Jaya AM. The effectiveness of giving plant PGPR rhizosphere bamboo on cocoa seeds germination at the nursery level. BIOMEDNATPROCH. 2021;10:1–5.
Khadeejath Rajeela TH, Gopal M, Gupta A, Bhat R, Thomas GV. Cross-compatibility evaluation of plant growth promoting rhizobacteria of coconut and cocoa on yield and rhizosphere properties of vegetable crops. Biocatal Agric Biotechnol. 2017;9:67–73.
Ferrás NegrÃn Y, Bustamante González CA, OrtÃz Gómez N. RAC. 2022. https://doi.org/10.15517/rac.v46i1.49873. Pseudomona sp en la emergencia de semillas y el desarrollo de posturas de cacao.
Beck E, Ziegler P. Biosynthesis and degradation of starch in higher plants. Annu Rev Plant Physiol Plant Mol Biol. 1989;40:95–117.
Carrillo A, Li C, Bashan Y. Increased acidification in the rhizosphere of cactus seedlings induced by Azospirillum brasilense. Naturwissenschaften. 2002;89:428–32.
Malik DK, Sindhu SS. Production of indole acetic acid by Pseudomonas sp.: effect of coinoculation with Mesorhizobium sp. Cicer on nodulation and plant growth of Chickpea (Cicer arietinum). Physiol Mol Biol Plants. 2011;17:25–32.
Chen B, Luo S, Wu Y, Ye J, Wang Q, Xu X, et al. The effects of the endophytic bacterium Pseudomonas fluorescens Sasm05 and IAA on the plant growth and cadmium uptake of sedum Alfredii hance. Front Microbiol. 2017;8:2538.
Suresh P, Vellasamy S, Ramamoorthy V. Implication of indole acetic acid and biofilm production by Pseudomonas fluorescens during tomato (Solanum lycopersicum) plant interactions. Biocatal Agric Biotechnol. 2025;65: 103554.
Alsultan W, Vadamalai G, Khairulmazmi A, Saud HM, Al-Sadi AM, Rashed O, et al. Isolation, identification and characterization of endophytic bacteria antagonistic to Phytophthora palmivora causing black pod of cocoa in Malaysia. Eur J Plant Pathol. 2019;155:1077–91.
Stephen L-K, Richard TA, Fredrick K. Biological control of black pod disease of cocoa (Theobroma Cacao L.) with Bacillus amyloliquefaciens, Aspergillus sp. and Penicillium sp. in vitro and in the field. J Microbiol Antimicrob. 2020;12:52–63.
Haas D, Défago G. Biological control of soil-borne pathogens by fluorescent pseudomonads. Nat Rev Microbiol. 2005;3:307–19.
Van Loon LC, Bakker PAHM, Pieterse CMJ. Systemic resistance induced by rhizosphere bacteria. Annu Rev Phytopathol. 1998;36:453–83.
Pieterse CMJ, Zamioudis C, Berendsen RL, Weller DM, Van Wees SCM, Bakker PAHM. Induced systemic resistance by beneficial microbes. Annu Rev Phytopathol. 2014;52:347–75.
Kumar A, Kumar SPJ, Chintagunta AD, Agarwal DK, Pal G, Singh AN, et al. Biocontrol potential of Pseudomonas stutzeri endophyte from Withania somnifera (Ashwagandha) seed extract against pathogenic Fusarium oxysporum and Rhizoctonia Solani. Arch Phytopathol Plant Prot. 2022;55:1–18.
Research Institute for Biotechnology and Environment, University NL, City HCM, Vietnam, Tran VT, Nguyen HT, Nguyen HT, Le DD, Faculty of biological sciences, Nong lam university, Ho Chi Minh city, vietnam. Isolation and characteristics of Pseudomonas fluorescens to inhibit Phytophthora palmivora causing rot disease in Durian. J Agric Dev. 2023;22:31–8.
Barahona E, Navazo A, MartÃnez-Granero F, Zea-Bonilla T, Pérez-Jiménez RM, MartÃn M, et al. Pseudomonas fluorescens F113 mutant with enhanced competitive colonization ability and improved biocontrol activity against fungal root pathogens. Appl Environ Microbiol. 2011;77:5412–9.
Vivekananthan R, Ravi M, Ramanathan A, Samiyappan R. Lytic enzymes induced by Pseudomonas fluorescens and other biocontrol organisms mediate defence against the anthracnose pathogen in Mango. World J Microbiol Biotechnol. 2004;20:235–44.