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Ayumu the chimpanzee spontaneously produced long, multicomponent instrumental displays by drumming, dragging, and throwing self-detached objects. Transition and rhythm analyses revealed non-random sequencing partially resembling pant-hoot structure, predominantly isochronous timing, and a more stable tempo when using tools than with the body. Accompanying play-face and silent bared teeth expressions suggest high arousal and positive affect, supporting the idea that affective vocal expression can be externalized through instrumental sound. Image credit: Hattori et al., doi: 10.1111/nyas.70239.

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