Toby Houlton won the wildlife behaviour category and overall Scottish Nature Photographer of the Year award with his image of tiny flies.
His picture, Dancing Gnats, was taken at Little Garve in Ross-shire.
“I had been walking back beside the river in the late afternoon, and there, in patches of dappled sun, the gnats were ‘dancing’,” said Toby.
“A moment of inspiration made me realise that taking an image with a slow shutter speed would create the amazing light trails, and the hint of the darker woodland behind would provide the sense of place.”
He added: “The speed of these tiny dancers meant that multiple wingbeats were captured in the exposure, creating the patterns.”
Competition judge Niall Irvine praised Toby for his creative use of photographic technique.