Villaseñor, A., Bobe, R. & Behrensmeyer, A. K. Middle Pliocene hominin distribution patterns in Eastern Africa. J. Hum. Evol. 147, 102856 (2020).

Article 
PubMed 

Google Scholar
 

Wood, B. & Strait, D. Patterns of resource use in early Homo and Paranthropus. J. Hum. Evolution 46, 119–162 (2004).

Article 

Google Scholar
 

Wood, B. & Boyle, E. K. Hominin taxic diversity: fact or fantasy? Am. J. Phys. Anthropol. 159, S37–S78 (2016).

Article 
PubMed 

Google Scholar
 

Haile-Selassie, Y., Suwa, G. & White, T. D. Late Miocene teeth from middle Awash, Ethiopia, and early hominid dental evolution. Science 303, 1503–1505 (2004).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

White, T. D. et al. Ardipithecus ramidus and the paleobiology of early hominids. Science 326, 64–86 (2009).

Article 
ADS 

Google Scholar
 

White, T. D. et al. Asa Issie, Aramis and the origin of Australopithecus. Nature 440, 883–889 (2006).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

Haile-Selassie, Y., Melillo, S. M. & Su, D. F. The Pliocene hominin diversity conundrum: do more fossils mean less clarity? Proc. Natl Acad. Sci. USA 113, 6364–6371 (2016).

Article 
ADS 
CAS 
PubMed 
PubMed Central 

Google Scholar
 

Alemseged, Z. et al. A juvenile early hominin skeleton from Dikika, Ethiopia. Nature 443, 296–301 (2006).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

Wynn, J. G. et al. Geological and palaeontological context of a Pliocene juvenile hominin at Dikika, Ethiopia. Nature 443, 332–336 (2006).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

White, T. D., Suwa, G., Simpson, S. & Asfaw, B. Jaws and teeth of Australopithecus afarensis from Maka, Middle Awash, Ethiopia. Am. J. Phys. Anthropol. 111, 45–68 (2000).

Article 
CAS 
PubMed 

Google Scholar
 

Kimbel, W. H. & Delezene, L. K. “Lucy” redux: a review of research on Australopithecus afarensis. Yearb. Phys. Anthropol. 52, 2–48 (2009).

Article 

Google Scholar
 

Asfaw, B. et al. Australopithecus garhi: a new species of early hominid from Ethiopia. Science 284, 629–635 (1999).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

McPherron, S. P. et al. Evidence for stone-tool-assisted consumption of animal tissues before 3.39 million years ago at Dikika, Ethiopia. Nature 466, 857–860 (2010).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

Villmoare, B. et al. Early Homo at 2.8 Ma from Ledi-Geraru, Afar, Ethiopia. Science 347, 1352–1355 (2015).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

Villmoare, B. et al. New discoveries of Australopithecus and Homo from Ledi-Geraru, Ethiopia. Nature https://doi.org/10.1038/s41586-025-09390-4 (2025).

Semaw, S. et al. 2.5-million-year-old stone tools from Gona, Ethiopia. Nature 385, 333–336 (1997).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

Braun, D. R. et al. Earliest known Oldowan artifacts at >2.58 Ma from Ledi-Geraru, Ethiopia, highlight early technological diversity. Proc. Natl Acad. Sci. USA 116, 11714–11717 (2019).

Article 
ADS 

Google Scholar
 

Alemseged, Z. et al. Fossils from Mille-Logya, Afar, Ethiopia, elucidate the link between Pliocene environmental changes and Homo origins. Nat. Commun. 11, 1–12 (2020).

Article 

Google Scholar
 

Kimbel, W. H., Johanson, D. C. & Rak, Y. Systematic assessment of a maxilla of Homo from Hadar, Ethiopia. Am. J. Phys. Anthropol. 103, 235–262 (1997).

Article 
CAS 
PubMed 

Google Scholar
 

Asfaw, B. et al. Remains of Homo erectus from Bouri, Middle Awash, Ethiopia. Nature 416, 317–320 (2002).

Article 
ADS 
PubMed 

Google Scholar
 

Kalb, J. E. et al. Fossil mammals and artefacts from the Middle Awash Valley, Ethiopia. Nature 298, 25–29 (1982).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

White, T. D. et al. Pleistocene Homo sapiens from Middle Awash, Ethiopia. Nature 423, 742–747 (2003).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

Broom, R. The Pleistocene anthropoid apes of South Africa. Nature 142, 377–379 (1938).

Article 
ADS 

Google Scholar
 

Grine, F. E. in Evolutionary History of the Robust Australopithecines (ed. Grine, F. E.) Ch. 30 (Routledge, 1988).

Robinson, J. T. The genera and species of the Australopithecinae. Am. J. Phys. Anthropol. 12, 181–200 (1954).

Article 
CAS 
PubMed 

Google Scholar
 

Wood, B. & Constantino, P. Paranthropus boisei: fifty years of evidence and analysis. Yearb. Phys. Anthropol. 134, 106–132 (2007).

Article 

Google Scholar
 

Plummer, T. W. et al. Expanded geographic distribution and dietary strategies of the earliest Oldowan hominins and Paranthropus. Science 379, 561–566 (2023).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

Harrison, T. in Paleontology and Geology of Laetoli: Human Evolution in Context vol. 2 (ed. Harrison, T.) Ch. 7 (Springer, 2011).

Suwa, G. et al. The first skull of Australopithecus boisei. Nature 389, 489–492 (1997).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

White, T. D., Asfaw, B. & Suwa, G. Pliocene hominid fossils from Gamedah, Middle Awash, Ethiopia. Trans. R. Soc. South Afr. 60, 79–83 (2005).

Article 

Google Scholar
 

Geraads, D. et al. Plio-Pleistocene mammals from Mille-Logya, Ethiopia, and the post-Hadar faunal change. J. Quat. Sci. 36, 1073–1089 (2021).

Article 

Google Scholar
 

deMenocal, P. B. African climate change and faunal evolution during the Pliocene–Pleistocene. Earth Planet. Sc. Lett. 220, 3–24 (2004).

Article 
ADS 
CAS 

Google Scholar
 

Bobe, R. & Eck, G. G. Responses of African bovids to Pliocene climatic change. Paleobiology 27, 1–47 (2001).

Article 

Google Scholar
 

Wynn, J. G. et al. in The Geology of Early Humans in the Horn of Africa (eds Quade, J. & Wynn, J. G.) Ch. 4 (Geological Society of America, 2008).

Levin, N. E. et al. in African Paleoecology and Human Evolution vol. 2 (eds Reynolds, S. C. & Bobe, R.) Ch. 17 (Cambridge Univ. Press, 2022).

Cerling, T. E. et al. Stable isotope-based diet reconstructions of Turkana Basin hominins. Proc. Natl Acad. Sci. USA 110, 10501–10506 (2013).

Article 
ADS 
CAS 
PubMed 
PubMed Central 

Google Scholar
 

Sponheimer, M. et al. Isotopic evidence of early hominin diets. Proc. Natl Acad. Sci. USA 110, 10513–10518 (2013).

Article 
ADS 
CAS 
PubMed Central 

Google Scholar
 

Negash, E. W. et al. Modern African ecosystems as landscape-scale analogues for reconstructing woody cover and early hominin environments. J. Hum. Evol. 197, 103604 (2024).

Article 
PubMed 

Google Scholar
 

Rak, Y., Kimbel, W. H., Moggi-Cecchi, J., Lockwood, C. A. & Menter, C. The DNH 7 skull of Australopithecus robustus from Drimolen (Main Quarry), South Africa. J. Hum. Evol. 151, 102913 (2021).

Article 
PubMed 

Google Scholar
 

Wood, B. A., Abbott, S. A. & Uytterschaut, H. Analysis of the dental morphology of Plio-Pleistocene hominids. IV. Mandibular postcanine root morphology. J. Anat. 156, 107–139 (1988).

CAS 
PubMed 
PubMed Central 

Google Scholar
 

Moore, N. C., Thackeray, J. F., Hublin, J.-J. & Skinner, M. M. Premolar root and canal variation in South African Plio-Pleistocene specimens attributed to Australopithecus africanus and Paranthropus robustus. J. Hum. Evol. 93, 46–62 (2016).

Article 
PubMed 

Google Scholar
 

Martin, L. Significance of enamel thickness in hominoid evolution. Nature 314, 260–263 (1985).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

Skinner, M. M., Alemseged, Z., Gaunitz, C. & Hublin, J.-J. Enamel thickness trends in Plio-Pleistocene hominin mandibular molars. J. Hum. Evol. 85, 35–45 (2015).

Article 
PubMed 

Google Scholar
 

Harrison, T. in Human Paleontology and Prehistory (eds Marom, A. & Hovers, E.) Ch. 4 (Springer, 2017).

Quinn, R. L. & Lepre, C. J. Contracting eastern African C4 grasslands during the extinction of Paranthropus boisei. Sci. Rep. 11, 7164 (2021).

Article 
ADS 
CAS 
PubMed 
PubMed Central 

Google Scholar
 

O’Brien, K., Hebdon, N. & Faith, J. T. Paleoecological evidence for environmental specialization in Paranthropus boisei compared to early Homo. J. Hum. Evol. 177, 103325 (2023).

Article 
PubMed 

Google Scholar
 

Suwa, G., White, T. D. & Howell, F. C. Mandibular postcanine dentition from the Shungura Formation, Ethiopia: crown morphology, taxonomic allocations, and Plio-Pleistocene hominid evolution. Am. J. Phys. Anthropol. 101, 247–282 (1996).

Article 
CAS 
PubMed 

Google Scholar
 

Deino, A. L. in Paleontology and Geology of Laetoli: Human Evolution in Context vol. 1 (ed. Harrison, T.) Ch. 4 (Springer, 2011).

Feibel, C. S., Brown, F. H. & McDougall, I. Stratigraphic context of fossil hominids from the Omo group deposits: Northern Turkana Basin, Kenya and Ethiopia. Am. J. Phys. Anthropol. 78, 595–622 (1989).

Article 
CAS 
PubMed 

Google Scholar
 

Arambourg, C. & Coppens, Y. Découverte d’un Australopithécien nouveau dans les gisements de l’Omo. S. Afr. J. Sci. 64, 58–59 (1968).

White, T. D. & Johanson, D. C. Pliocene hominid mandibles from the Hadar Formation, Ethiopia: 1974-1977 collections. Am. J. Phys. Anthropol. 57, 501–544 (1982).

Article 

Google Scholar
 

White, T. D. New fossil hominids from Laetolil, Tanzania. Am. J. Phys. Anthropol. 46, 197–230 (1977).

Article 
CAS 
PubMed 

Google Scholar
 

White, T. D. Additional fossil hominids from Laetoli, Tanzania: 1976–1979 specimens. Am. J. Phys. Anthropol. 53, 487–504 (1980).

Article 

Google Scholar
 

Kimbel, W. H., Rak, Y. & Johanson, D. C. in The Skull of Australopithecus afarensis (eds. Ciochon, R. & Wood, B.) 202–203 (Oxford Univ. Press, 2004).

Alemseged, Z. et al. A new hominin from the Basal Member of the Hadar Formation, Dikika, Ethiopia, and its geological context. J. Hum. Evol. 49, 499–514 (2005).

Article 
PubMed 

Google Scholar
 

Haile-Selassie, Y. & Ryan, T. M. Comparative description and taxonomy of new hominin juvenile mandibles from the Pliocene of Woranso-Mille (Central Afar, Ethiopia). J. Hum. Evol. 132, 15–31 (2019).

Article 
PubMed 

Google Scholar
 

Melillo, S. M. et al. New Pliocene hominin remains from the Leado Dido’a area of Woranso-Mille, Ethiopia. J. Hum. Evol. 153, 102956 (2021).

Article 
PubMed 

Google Scholar
 

Coppens, Y. Les restes d’Hominidés des séries inférieures et moyennes des formations plio-villafranchiennes de l’Omo en Ethiopie. C. R. Acad. Sci. III 271, 2286–2289 (1970).


Google Scholar
 

Leakey, R. E. F. & Walker, A. New Australopithecus boisei specimens from east and west Lake Turkana, Kenya. Am. J. Phys. Anthropol. 76, 1–24 (1988).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

Suwa, G. A Comparative Analysis of Hominid Dental Remains from the Shungura and Usno Formations, Omo Valley, Ethiopia. PhD thesis, Univ. of California at Berkeley (1990).

Wood, B. A. Koobi Fora Research Project: Volume 4: Hominid Cranial Remains (Oxford Univ. Press, 1991).

Howell, F. C. Remains of hominidae from Pliocene/Pleistocene formations in the lower Omo basin, Ethiopia. Nature 223, 1234–1239 (1969).

Article 
ADS 
CAS 
PubMed 

Google Scholar
 

Coppens, Y. Les restes d’Hominidés des séries inférieures des formations plio-villafranchiennes de l’Omo en Ethiopie. C. R. Acad. Sci. III 272, 36–39 (1970).


Google Scholar
 

Brown, B., Brown, F. H. & Walker, A. New hominids from the Lake Turkana Basin, Kenya. J. Hum. Evol. 41, 29–44 (2001).

Article 
CAS 
PubMed 

Google Scholar
 

Prat, S., Brugal, J.-P., Roche, H. & Texier, P.-J. Nouvelles découvertes de dents d’hominidés dans le membre Kaitio de la formation de Nachukui (1,65–1,9 Ma), Ouest du lac Turkana (Kenya). C. R. Palevol. 2, 685–693 (2003).

Article 

Google Scholar
 

Bromage, T., Schrenk, F. & Juwayeyi, Y. Paleobiogeography of the Malawi Rift: age and vertebrate paleontology of the Chiwondo Beds, northern Malawi. J. Hum. Evol. 28, 37–57 (1995).

Article 

Google Scholar
 

Grine, F. E. New hominid fossils from the Swartkrans Formation (1979–1986 excavations): craniodental specimens. Am. J. Phys. Anthr. 79, 409–449 (1989).

Article 
CAS 

Google Scholar
 

Grine, F. E. & Strait, D. S. New hominid fossils from member 1 “Hanging Remnant”, Swartkrans formation, South Africa. J. Hum. Evol. 26, 57–75 (1994).

Article 

Google Scholar
 

Moggi-Cecchi, J., Menter, C., Boccone, S. & Keyser, A. Early hominin dental remains from the Plio-Pleistocene site of Drimolen, South Africa. J. Hum. Evol. 58, 374–405 (2010).

Article 
PubMed 

Google Scholar
 

Pickering, T. R. et al. New early Pleistocene hominin teeth from the Swartkrans formation, South Africa. J. Hum. Evol. 100, 1–15 (2016).

Article 
PubMed 

Google Scholar
 

Leece, A. B. et al. New hominin dental remains from the Drimolen Main Quarry, South Africa (1999–2008). Am. J. Biol. Anthr. 179, 240–260 (2022).


Google Scholar
 

Leece, A. B. et al. New hominin dental remains from the ∼2.04–1.95 Ma Drimolen Main Quarry, South Africa. Ann. Hum. Biol. 50, 407–427 (2023).

Article 
CAS 
PubMed 

Google Scholar
 

Braga, J. & Grine, F. E. New craniodental fossils of Paranthropus robustus from Kromdraai, South Africa (2014–2017 excavations). J. Hum. Evol. 188, 103481 (2024).

Article 
PubMed 

Google Scholar
 

Haile-Selassie, Y. et al. Dentognathic remains of Australopithecus afarensis from Nefuraytu (Woranso-Mille, Ethiopia): comparative description, geology, and paleoecological context. J. Hum. Evol. 100, 35–53 (2016).

Article 
PubMed 

Google Scholar
 

Hanegraef, H. & Spoor, F. Morphological variation of the Australopithecus afarensis maxilla. J. Hum. Evol. 201, 103651 (2025).

Article 
PubMed 

Google Scholar