Hussain, S. Z., Naseer, B., Qadri, T., Fatima, T. & Bhat, T. A. Walnut (Juglans regia)- Morphology, Taxonomy, Composition and Health Benefits. . in Fruits Grown in Highland Regions of the Himalayas 269–281 (2021).
Bhardwaj, A., Singh, A., Patnaik, R. S. & Bhardwaj, S. Juglans regia L: A review of its traditional uses phytochemistry and therapeutic applications. J. Pharm. Negat. Results. 14, 11–16 (2023).
Hayes, D., Angove, M. J., Tucci, J. & Dennis, C. Walnuts (Juglans regia) chemical composition and research in human health. Crit. Rev. Food Sci. Nutr. 56, 1231–1241 (2016).
Tapia, M. I. et al. Comparative study of the nutritional and bioactive compounds content of four walnut (Juglans regia L.) cultivars. J. Food Compos. Anal. 31, 232–237 (2013).
Jahanbani, R. et al. Antioxidant and anticancer activities of walnut (Juglans regia L.) protein hydrolysates using different proteases. Plant Foods Hum. Nutr. 71, 402–409 (2016).
Woeste, K. Thirty polymorphic nuclear microsatellite loci from black walnut. J. Hered. 93, 58–60 (2002).
Victory, E. R., Glaubitz, J. C., Rhodes, O. E. & Woeste, K. E. Genetic homogeneity in Juglans Nigra (Juglandaceae) at nuclear microsatellites. Am. J. Bot. 93, 118–126 (2006).
Aradhya, M. et al. Genetic and ecological insights into glacial refugia of walnut (Juglans regia L). PLoS One. 12, e0185974 (2017).
Pollegioni, P. et al. Biocultural diversity of common walnut (Juglans regia L.) and sweet chestnut (Castanea sativa Mill.) across Eurasia. Ecol. Evol. 10, 11192–11216 (2020).
FAO. Crops and livestock products. FAOSTAT Crops and livestock products https://www.fao.org/faostat/en/#data/QCL (2024).
TÜİK. Türkiye İstatistik Kurumu. Preprint at. (2022).
Kadakoğlu, B., Bayav, A. & Karli, B. Türkiye’de Ceviz üretim Projeksiyonu ve Rekabet Gücü analizi. Meyve Bilimi. 9, 8–15 (2022).
Bernard, A., Lheureux, F. & Dirlewanger, E. Walnut: past and future of genetic improvement. Tree Genet. Genomes. 14, 1 (2018).
Orhan, E. et al. Genetic diversity detection of seed-propagated walnut (Juglans regia L.) germplasm from Eastern Anatolia using SSR markers. Folia Horticulturae. 32, 37–46 (2020).
Bozhuyuk, M. R. & Walnut Fruit, A great morphological, biochemical and molecular biodiversity in Turkey and in the world. In Integrated Alternative Farming Models Vol. 211 (eds Kokten, K. & Özdemir, S.) 211–259 (Ankara, 2022).
Adigüzel, P. & Solmaz, İ. Türkiye’de Bitki genetik Kaynaklarının Mevcut Durumu ve Korunması. Turk. Tarim Arast Derg. 10, 352–360 (2023).
Magige, E. A. et al. Genetic diversity and structure of Persian walnut (Juglans regia L.) in pakistan: implications for conservation. Plants 11, 1652 (2022).
Chesnokov, Y. V., Kosolapov, V. M. & Savchenko, I. V. Morphological genetic markers in plants. Russ J. Genet. 56, 1406–1415 (2020).
Schulman, A. H. Molecular markers to assess genetic diversity. Euphytica 158, 313–321 (2007).
Budak, H., Shearman, R. C., Parmaksiz, I. & Dweikat, I. Comparative analysis of seeded and vegetative biotype buffalograsses based on phylogenetic relationship using issrs, ssrs, rapds, and SRAPs. Theor. Appl. Genet. 109, 280–288 (2004).
Kabiri, G., Bouda, S. & Haddioui, A. Evaluation of genetic diversity and structuration across altitude of walnut (Juglans regia L.) accessions from Morocco using SSR markers. Bulgarian J. Agricultural Sci. 28, 451–458 (2022).
Shah, R. A. et al. Morphological to Molecular Markers: Plant Genetic Diversity Studies in Walnut (Juglans regia L.)—A Review. . Erwerbs-Obstbau 65, 1499–1511 (2023).
Potts, S. M. et al. Genetic diversity and characterization of a core collection of Malus germplasm using simple sequence repeats (SSRs). Plant. Mol. Biol. Rep. 30, 827–837 (2012).
Fan, L. et al. Transferability of newly developed Pear SSR markers to other rosaceae species. Plant. Mol. Biol. Rep. 31, 1271–1282 (2013).
Başak, İ., Muradoğlu, F., Özer, G. & Güler, E. Comprehensive Genetic Diversity Assessment of Promising Walnut (Juglans regia L.) Genotypes Using RAPD, ISSR, and iPBS Markers. Preprint at (2024). https://doi.org/10.21203/rs.3.rs-4946835/v1
Shah, U. N., Mir, J., Ahmed, N. & Fazili, K. M. Genetic Diversity Analysis of Walnut (Juglans regia L.) from Kashmir Valley Using RAPD and ISSR Markers. Agrotechnology 08, (2019).
Kabiri, G., Bouda, S., Elhansali, M. & Haddioui, A. Genetic diversity and structure of walnut (Juglans regia L.) genotypes from middle and high atlas mountains of Morocco as investigated by Inter-Simple sequence repeat (ISSR) markers. Aust J. Crop Sci. 13, 1983–1991 (2019).
Christopoulos, M. V., Rouskas, D., Tsantili, E. & Bebeli, P. J. Germplasm diversity and genetic relationships among walnut (Juglans regia L.) cultivars and Greek local selections revealed by Inter-Simple sequence repeat (ISSR) markers. Sci. Hortic. 125, 584–592 (2010).
Ji, A. et al. Genetic diversity and population structure of North China mountain walnut revealed by ISSR. Am. J. Plant. Sci. 05, 3194–3202 (2014).
Ghanbari, A. et al. Genetic diversity evaluation of some walnut (Juglans regia L.) genotypes in Meshkin-Shahr by ISSR marker. J. Nuts. 10, 1–8 (2019).
Sharifi, S., Amiri-Fahliani, R., Kavoosi, B. & Asl, A. M. Walnut genetic diversity investigation using phenological and morphological characteristics and Issr markers. J. Agricultural Sci. Technol. 23, 187–200 (2021).
Bernard, A., Barreneche, T., Lheureux, F. & Dirlewanger, E. Analysis of genetic diversity and structure in a worldwide walnut (Juglans regia L.) germplasm using SSR markers. PLoS One. 13, e0208021 (2018).
Guney, M. et al. Genetic diversity among some walnut (Juglans regia L.) genotypes by SSR markers. Sustainability 13, 6830 (2021).
Nicese, F. P., Hormaza, J. I. & McGranahan, G. H. Molecular characterization and genetic relatedness among walnut (Juglans regia L.) genotypes based on RAPD markers. Euphytica 101, 199–206 (1998).
Xu, Z., Hu, T. & Zhang, F. Genetic diversity of walnut revealed by AFLP and RAPD markers. J. Agric. Sci. 4, 271–276 (2012).
Ahmed, N. et al. SSR and RAPD analysis of genetic diversity in walnut (Juglans regia L.) genotypes from Jammu and Kashmir, India. Physiology Mol. Biology Plants 18(2), 149–160 (2012).
Nadeem, M. A. et al. DNA molecular markers in plant breeding: current status and recent advancements in genomic selection and genome editing. Biotechnol. Biotechnol. Equip. 32, 261–285 (2018).
Sun, L. et al. Modern plant breeding techniques in crop improvement and genetic diversity: from molecular markers and gene editing to artificial Intelligence—A. Crit. Rev. Plants. 13, 2676 (2024).
Sevindik, E., Okan, K., Sevindik, M. & Ercisli, S. Genetic diversity and phylogenetic analyses of Juglans regia L. (Juglandaceae) populations using RAPD, ISSR markers and NrDNA ITS regions. Erwerbs-Obstbau 65, 311–320 (2023).
Xie, Q. et al. Assessment of genetic diversity and population structure of common walnut (Juglans regia) germplasm with simple sequence repeat (SSR) markers. Genet. Resour. Crop Evol. 72, 2265–2276 (2025).
Sütyemez, M., Özcan, A., Yılmaz, A., Yıldırım, E. & Bükücü, Ş. B. Determining phenological and genetic variation in genotypes obtained from open-pollinated seeds of ‘maraş 12’ walnut (Juglans regia L.) cultivar. Genet. Resour. Crop Evol. 69, 823–838 (2022).
Doyle, J. J. Isolation of plant DNA from fresh tissue. Focus (Madison) 12, 13–15 (1990).
Rasul, K. S. & Grundler, F. M. W. Abdul-razzak tahir, N. Genetic diversity and population structure assessment of Iraqi tomato accessions using fruit characteristics and molecular markers. Hortic. Environ. Biotechnol. 63, 523–538 (2022).
Yildiz, A., Hajyzadeh, M., Ozbek, K., Nadeem, M. A. & Hunter, D. Molecular characterisation of the oldest domesticated Turkish Einkorn wheat landraces with simple sequence repeat (SSR) markers. Biotechnol. Biotechnol. Equip. 35, 1291–1300 (2021).
Yeh, F. C., Yang, R., Boyle, T. J., Ye, Z. & Xiyan, J. M. PopGene32, Microsoft Windows-based freeware for population genetic analysis, version 1.32. Molecular Biology and Biotechnology Centre, University of Alberta, Edmonton, Alberta, Canada (2000).
Roldán-Ruiz, I., Dendauw, J., Van Bockstaele, E., Depicker, A. & De Loose M. AFLP markers reveal high polymorphic rates in ryegrasses (Lolium spp). Mol. Breeding. 6, 125–134 (2000).
Kovach, W. L. Mutivariate statistical package (MVSP) plus version 3.22 user’s manual. Publish Kovach Comput. Services 137, (2007).
Çilesiz, Y. et al. Identification of phenotypic diversity and DArTseq loci associated with vitamin A contents in Turkish common bean germplasm through GWAS. Plants 14, 776 (2025).