Meyer, M. et al. Nuclear DNA sequences from the Middle Pleistocene Sima de los Huesos hominins. Nature 531, 504\u2013507 (2016).<\/p>\n
Article<\/a>\u00a0 Duval, M. et al. The first direct ESR dating of a hominin tooth from Atapuerca Gran Dolina TD-6 (Spain) supports the antiquity of Homo antecessor. Quat. Geochronol. 47, 120\u2013137 (2018).<\/p>\n Article<\/a>\u00a0 Berm\u00fadez de Castro, J. M. et al. A hominid from the Lower Pleistocene of Atapuerca, Spain: possible ancestor to Neandertals and modern humans. Science 276, 13921395 (1997).<\/p>\n Article<\/a>\u00a0 Hublin, J.-J. et al. New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens. Nature 546, 289\u2013292 (2017).<\/p>\n Article<\/a>\u00a0 Rightmire, G. P. Human evolution in the Middle Pleistocene: the role of Homo heidelbergensis. Evol. Anthropol. 6, 218\u2013227 (1998).<\/p>\n Article<\/a>\u00a0 Mounier, A. & Caparros, M. The phylogenetic status of Homo heidelbergensis\u2014a cladistic study of Middle Pleistocene hominins. Bull. M\u00e9m. Soc. Anthop. Paris 27, 125 (2015).<\/p>\n Berm\u00fadez de Castro, J. M. & Martin\u00f3n-Torres, M. The origin of the Homo sapiens lineage: when and where? Quat. Int. 634, 1\u201313 (2022).<\/p>\n Article<\/a>\u00a0 Clarke, R. J. The Ndutu cranium and the origin of Homo sapiens. J. Hum. Evol. 19, 699\u2013736 (1990).<\/p>\n Article<\/a>\u00a0 Texier, J.-P. et al. Lithostratigraphy of the littoral deposits of the last one million years in the Casablanca region (Maroc). Quaternaire 13, 23\u201341 (2002).<\/p>\n Article<\/a>\u00a0 Lef\u00e8vre, D. et al. Les pal\u00e9olittoraux plio-pl\u00e9istoc\u00e8nes de Casablanca, cadre chronostratigraphique et pal\u00e9og\u00e9ographique de la Pr\u00e9histoire ancienne du Maroc atlantique. Bull. Arch\u00e9ol. Maroc. 21, 39\u201370 (2021).<\/p>\n Raynal, J.-P. & Texier, J.-P. D\u00e9couverte d\u2019Acheul\u00e9en ancien dans la carri\u00e8re Thomas 1 \u00e0 Casablanca et probl\u00e8me de l\u2019anciennet\u00e9 de la pr\u00e9sence humaine au Maroc. C. R. Acad. Sci. Paris 308, 1743\u20131749 (1989).<\/p>\n Raynal, J. P., Magoga, L., Sbihi-Alaoui, F.-Z. & Geraads, D. in The Earliest Occupation of Europe Vol. 27: Analecta Praehistorica Leidensia (eds Roebroeks, W. & Van Kolfschoten, T.) Ch. 13, 255\u2013262 (Univ. Leiden, 1995).<\/p>\n Gallotti, R. et al. First high resolution chronostratigraphy for the early North African Acheulean at Casablanca (Morocco). Sci. Rep. 11, 15340 (2021).<\/p>\n Article<\/a>\u00a0 Sausse, F. La mandibule atlanthropienne de la carri\u00e8re Thomas I (Casablanca). L\u2019Anthropologie 79, 81\u2013112 (1975).<\/p>\n Geraads, D., Beriro, P. & Roche, H. La faune et l\u2019industrie des sites \u00e0 Homo erectus des Carri\u00e8res Thomas (Maroc). Pr\u00e9cisions sur l’\u00e2ge de ces Hominid\u00e9s. C. R. Acad. Sci. Paris 291, 195\u2013198 (1980).<\/p>\n Ennouchi, E. D\u00e9couverte d\u2019un Pith\u00e9canthropien au Maroc. C. R. Acad. Sci. Paris 269, 763\u2013765 (1969).<\/p>\n Raynal, J.-P. et al. Hominid cave at Thomas Quarry I (Casablanca, Morocco): recent findings and their context. Quat. Int. 223-224, 369\u2013382 (2010).<\/p>\n Article<\/a>\u00a0 Raynal, J.-P. et al. in Handbook of Pleistocene Archaeology of Africa: Hominin Behavior, Geography, and Chronology (eds Beyin, A. et al.) 841\u2013861 (Springer, 2023).<\/p>\n Raynal, J. P. et al. Contextes et \u00e2ge des nouveaux restes dentaires humains du Pl\u00e9istoc\u00e8ne moyen de la carri\u00e8re Thomas I \u00e0 Casablanca (Maroc). Bull. Soc. Pr\u00e9hist. Fran\u00e7. 108, 645\u2013669 (2011).<\/p>\n Article<\/a>\u00a0 Geraads, D. et al. in African Paleoecology and Human Evolution (eds Bobe R. & Reynolds S. C.) 481\u2013491 (Cambridge Univ. Press, 2022).<\/p>\n Daujeard, C. et al. Pleistocene hominins as a resource for carnivores: a c. 500,000-year-old human femur bearing tooth-marks in North Africa (Thomas Quarry I, Morocco). PLoS ONE 11, e0152284 (2016).<\/p>\n Article<\/a>\u00a0 Channell, J. E. T., Singer, B. S. & Jicha, B. R. Timing of Quaternary geomagnetic reversals and excursions in volcanic and sedimentary archives. Quat. Sci. Rev. 228, 106114 (2020).<\/p>\n Article<\/a>\u00a0 Channell, J. E. T. Complexity in Matuyama\u2013Brunhes polarity transitions from North Atlantic IODP\/ODP deep-sea sites. Earth Planet. Sci. Lett. 467, 43\u201356 (2017).<\/p>\n Article<\/a>\u00a0 Haneda, Y., Okada, M., Suganuma, Y. & Kitamura, T. A full sequence of the Matuyama\u2013Brunhes geomagnetic reversal in the Chiba composite section, Central Japan. Prog. Earth Planet. Sci. 7, 44 (2020).<\/p>\n Article<\/a>\u00a0 Lisiecki, L. E. & Raymo, M. E. A Pliocene-Pleistocene stack of 57 globally distributed benthic \u03b418O records. Paleoceanography 20, PA1003 (2005).<\/p>\n ADS<\/a>\u00a0 Pickford, M. The fossil Suidae (Mammalia, Artiodactyla) from Ternifine (Tighenif) Algeria. M\u00fcnch. Geowiss. Abh. A 50, 1\u201367 (2020).<\/p>\n Geraads, D. Biochronologie mammalienne du Quaternaire du Maroc atlantique, dans son cadre r\u00e9gional. L\u2019Anthropologie 114, 324\u2013340 (2010).<\/p>\n Article<\/a>\u00a0 Larrasoa\u00f1a, J. C., Roberts, A. P. & Rohling, E. J. Dynamics of green Sahara periods and their role in hominin evolution. PLoS ONE 8, e76514 (2013).<\/p>\n Article<\/a>\u00a0 Larrasoa\u00f1a, J. C. A review of West African monsoon penetration during Green Sahara periods; implications for human evolution and dispersals over the last three million years. Oxford Open Climate Change 1, kgab011 (2021).<\/p>\n Article<\/a>\u00a0 Rhodes, E. J., Singarayer, J. S., Raynal, J. P., Westaway, K. E. & Sbihi-Alaoui, F. Z. New age estimates for the Palaeolithic assemblages and Pleistocene succession of Casablanca, Morocco. Quat. Sci. Rev. 25, 2569\u20132585 (2006).<\/p>\n Article<\/a>\u00a0 Gr\u00fcn, R. & Stringer, C. Direct dating of human fossils and the ever-changing story of human evolution. Quat. Sci. Rev. 322, 108379 (2023).<\/p>\n Article<\/a>\u00a0 Dobson, S. D. & Trinkaus, E. Cross-sectional geometry and morphology of the mandibular symphysis in Middle and Late Pleistocene Homo. J. Hum. Evol. 43, 67\u201387 (2002).<\/p>\n Article<\/a>\u00a0 Berm\u00fadez de Castro, J. M., Martin\u00f3n-Torres, M., G\u00f3mez-Robles, A. G., Prado, L. & Carbonell, E. New human evidence of the Early Pleistocene settlement of Europe, from Sima del Elefante site (Sierra de Atapuerca, Burgos, Spain). Quat. Int. 223-224, 431\u2013433 (2010).<\/p>\n Article<\/a>\u00a0 Rosas, A. & Berm\u00fadez de Castro, J. M. The ATD6-5 mandibular specimen from Gran Dolina (Atapuerca, Spain). Morphological study and phylogenetic implications. J. Hum. Evol. 37, 567\u2013590 (1999).<\/p>\n Article<\/a>\u00a0 Rosas, A. Occurrence of Neanderthal features in mandibles from the Atapuerca-SH site. Am. J. Phys. Anthropol. 114, 74\u201391 (2001).<\/p>\n Article<\/a>\u00a0 Vialet, A., Modesto-Mata, M., Martin\u00f3n-Torres, M., Mart\u00ednez de Pinillos, M. & Berm\u00fadez de Castro, J. M. A reassessment of the Montmaurin-La Niche mandible (Haute Garonne, France) in the context of European Pleistocene human evolution. PLoS ONE 13, e0189714 (2018).<\/p>\n Article<\/a>\u00a0 Mounier, A., Marchal, F. & Condemi, S. Is Homo heidelbergensis a distinct species? New insight on the Mauer mandible. J. Hum. Evol. 56, 219\u2013246 (2009).<\/p>\n Article<\/a>\u00a0 Rak, Y., Ginzburg, A. & Geffen, E. Does Homo neanderthalensis play a role in modern human ancestry? The mandibular evidence. Am. J. Phys. Anthropol. 119, 199204 (2002).<\/p>\n Article<\/a>\u00a0 Rosas, A. Seventeen new mandibular specimens from the Atapuerca\/Ibeas Middle Pleistocene Hominids sample (1985-1992). J. Hum. Evol. 28, 533\u2013559 (1995).<\/p>\n Article<\/a>\u00a0 Rosas, A. & Berm\u00fadez de Castro, J. M. The Mauer mandible and the evolutionary significance of Homo heidelbergensis. Geobios 31, 687\u2013697 (1998).<\/p>\n Article<\/a>\u00a0 Carbonell, E. et al. An Early Pleistocene hominin mandible from Atapuerca-TD6, Spain. Proc. Natl Acad. Sci. USA 102, 5674\u20135678 (2005).<\/p>\n Article<\/a>\u00a0 Berm\u00fadez de Castro, J. M., Martin\u00f3n-Torres, M., G\u00f3mez-Robles, A., Prado, L. & Sarmiento, S. Comparative analysis of the Gran-Dolina-TD6 (Spain) and Tighennif (Algeria) hominin mandibles. Bull. M\u00e9m. Soc. Anthropol. Paris 19, 166\u2013178 (2007).<\/p>\n Article<\/a>\u00a0 Boughner, J. C., Marchiori, D. F. & Packota, G. V. Unexpected variation of human molar size patterns. J. Hum. Evol. 161, 103072 (2021).<\/p>\n Article<\/a>\u00a0 Carretero, J. M., Lorenzo, C. & Arsuaga, J. L. Axial and appendicular skeleton of Homo antecessor. J. Hum. Evol. 37, 459\u2013499 (1999).<\/p>\n Article<\/a>\u00a0 Sahnouni, M. & van der Made, J. in The Cutting Edge: New Approaches to the Archaeology of Human Origins (eds Schick, K. & Toth, N.) 179\u2013210 (Stone Age Institute Press, 2009).<\/p>\n Geraads, D. in African Paleoecology and Human Evolution (eds Bobe, R. & Reynolds, S. C.) 475\u2013480 (Cambridge Univ. Press, 2022).<\/p>\n Geraads, D. Pleistocene Carnivora (Mammalia) from Tighennif (Ternifine), Algeria. Geobios 49, 445\u2013458 (2016).<\/p>\n Article<\/a>\u00a0 Martini, P. & Geraads, D. Camelus thomasi Pomel, 1893 from the Pleistocene typelocality Tighennif (Algeria). Comparisons with modern Camelus. Geodiversitas 40, 115\u2013134 (2018).<\/p>\n Article<\/a>\u00a0 Martin\u00f3n-Torres, M. et al. in Dental Perspectives on Human Evolution: State of the Art Research in Dental Paleoanthropology (eds Bailey, S. & Hublin, J.-J.) Ch. 5, 65\u201379 (Springer, 2007).<\/p>\n Berm\u00fadez de Castro, J. M. et al. Comparative dental study between Homo antecessor and Chinese Homo erectus: Nonmetric features and geometric morphometrics. J. Hum. Evol. 161, 103087 (2021).<\/p>\n Article<\/a>\u00a0 G\u00f3mez-Robles, A. Dental evolutionary rates and its implications for the Neanderthal modern human divergence. Sci. Adv. 5, eaaw1268 (2019).<\/p>\n Article<\/a>\u00a0 Feng, X. et al. The phylogenetic position of the Yunxian cranium elucidates the origin of Homo longi and the Denisovans. Science 389, 1320\u20131324 (2025).<\/p>\n Article<\/a>\u00a0 Harrison, R. J., & Feinberg, J. M. FORCinel: an improved algorithm for calculating first-order reversal curve distributions using locally weighted regression smoothing. Geochem. Geophys. Geosyst. https:\/\/doi.org\/10.1029\/2008GC001987<\/a> (2008).<\/p>\n Bergmann, I., Hublin, J.-J., Gunz, P. & Freidline, S. E. How did modern morphology evolve in the human mandible? The relationship between static adult allometry and mandibular variability in Homo sapiens. J. Hum. Evol. 157, 103026 (2021).<\/p>\n Article<\/a>\u00a0 Bergmann, I. et al. The relevance of late MSA mandibles on the emergence of modern morphology in Northern Africa. Sci. Rep. 12, 8841 (2022).<\/p>\n Article<\/a>\u00a0 Freidline, S. E., Gunz, P., Harvati, K. & Hublin, J.-J. Middle Pleistocene human facial morphology in an evolutionary and developmental context. J. Hum. Evol. 63, 723\u2013740 (2012).<\/p>\n Article<\/a>\u00a0 Freidline, S. E., Gunz, P., Harvati, K. & Hublin, J.-J. Evaluating developmental shape changes in Homo antecessor subadult facial morphology. J. Hum. Evol. 65, 404\u2013423 (2013).<\/p>\n Article<\/a>\u00a0 Freidline, S. E., Gunz, P., Jankovi\u0107, I., Harvati, K. & Hublin, J. J. A comprehensive morphometric analysis of the frontal and zygomatic bone of the Zuttiyeh fossil from Israel. J. Hum. Evol. 62, 225\u2013241 (2012).<\/p>\n Article<\/a>\u00a0 Freidline, S. E. et al. Early presence of Homo sapiens in Southeast Asia by 86-68 kyr at Tam Pa Ling, Northern Laos. Nat. Commun. 14, 3193 (2023).<\/p>\n Article<\/a>\u00a0 Wiley, D. F. et al. In Proc. 16th IEEE Visualization Conference (VIS 2005) Vol. 55, 431\u2013438 (IEEE, 2005).<\/p>\n R Core Team. R: A Language and Environment for Statistical Computing (R Foundation for Statistical Computing, 2021); www.R-project.org\/<\/a>.<\/p>\n Schlager, S. in Statistical Shape and Deformation Analysis (eds Zheng, S. L. G. & Szekely, G.) 217\u2013256 (Academic, 2017).<\/p>\n Adams, D. et al. Geomorph: software for geometric morphometric analyses. R package version 4.0.2 (2021); cran.rproject.org\/package=geomorph<\/a>.<\/p>\n Baken, E. K. et al. geomorph v4.0 and gmShiny: enhanced analytics and a new graphical interface for a comprehensive morphometric experience. Methods Ecol. Evol. 12, 2355\u20132363 (2021).<\/p>\n Article<\/a>\u00a0 Gunz, P. et al. Principles for the virtual reconstruction of hominin crania. J. Hum. Evol. 57, 48\u201362 (2009).<\/p>\n Article<\/a>\u00a0 Gunz, P., Mitteroecker P. & Bookstein F. in Modern Morphometrics in Physical Anthropology (ed. Slice, D. E.) 73\u201398 (Plenum, 2005).<\/p>\n Gunz, P. & Mitteroecker, P. Semilandmarks: a method for quantifying curves and surfaces. Hystrix 24, 103\u2013109 (2013).<\/p>\n Rohlf, F. J. & Slice, D. Extensions of the Procrustes method for the optimal superimposition of landmarks. Syst. Biol. 39, 40\u201359 (1990).<\/p>\n Arambourg, C. & Biberson, P. The fossil human remains from the Paleolithic site of Sidi Abderrahman (Morocco). Am. J. Phys. Anthropol. 14, 467\u2013489 (1956).<\/p>\n Article<\/a>\u00a0 Rightmire, G. P. Comparative studies of Late Pleistocene human remains from Klasies River Mouth, South Africa. J. Hum. Evol. 20, 131\u2013156 (1991).<\/p>\n Article<\/a>\u00a0 Stewart, T. D. in Annual Report of the Smithsonian Institution 521\u2013533 (US Government Printing Office, 1962).<\/p>\n Tillier, A.-M. in Le Squelette Moust\u00e9rien de K\u00e9bara 2 97\u2013112 (CNRS, 1991).<\/p>\n Stewart, T. D. The Neanderthal skeletal remains from Shanidar Cave, Iraq: a summary of findings to date. Proc. Am. Phil. Soc. 121, 121\u2013165 (1977).<\/p>\n Trinkaus, E. The Shanidar Neanderthals (Academic, 1983).<\/p>\n Leroi-Gourhan, A. \u00c9tude des Restes Humains Fossiles provenant des Grottes d\u2019Arcysur-Cure (Masson et Cie, 1958).<\/p>\n Daura, J. et al. A Neandertal mandible from the Cova del Gegant (Sitges, Barcelona, Spain). J. Hum. Evol. 49, 56\u201370 (2005).<\/p>\n Article<\/a>\u00a0 Topinard, P. Les caract\u00e8res simiens de la m\u00e2choire de la Naulette. Rev. Antropol. 15, 385\u2013431 (1886).<\/p>\n Blake, C. C. On a human jaw from the cave of La Naulette, near Dinant, Belgium. Anthropol. Rev. 5, 294\u2013303 (1867).<\/p>\n Article<\/a>\u00a0 Leguebe, A. & Toussaint, M. in La Mandibule et les Cubitus de la Naulette: Morphologie et Morphom\u00e9trie 15 (CNRS, 1988).<\/p>\n Heim, J. L. Les hommes fossiles de La Ferrassie (Dordogne) et le probl\u00e8me de la d\u00e9finition des Neandertaliens classiques. III. Squelette c\u00e9phalique. L\u2019Anthropologie 78, 321\u2013378 (1974).<\/p>\n de Lumley, M.-A. Les N\u00e9andertaliens de la grotte de l\u2019Hortus. Etud. Quat. 1, 375\u2013385 (1972).<\/p>\n Condemi, S. et al. Possible interbreeding in late Italian Neanderthals? New data from the Mezzena jaw (Monti Lessini, Verona, Italy). PLoS ONE 8, e59781 (2013).<\/p>\n Article<\/a>\u00a0 Corrain, C. Resti scheletrici umani del \u2018Riparo Mezzena\u2019. Mem. Mus. Civ. Stor. Nat. Verona 16, 97\u2013101 (1968).<\/p>\n Walker, M. J., Lombardi, A. V., Zapata, J. & Trinkaus, E. Neandertal mandibles from the Sima de las Palomas del Cabezo Gordo, Murcia, southeastern Spain. Am. J. Phys. Anthropol. 142, 261\u2013272 (2010).<\/p>\n Article<\/a>\u00a0 Martin, H. Machoire humaine moust\u00e9rienne trouv\u00e9e dans la station de La Quina. L\u2019homme Pr\u00e9hist. 13, 3\u201321 (1926).<\/p>\n Martin, H. Position stratigraphique des Ossements humains recueillis dans le Moust\u00e9rien de La Quina de 1908 \u00e0 1912. Bull. Soc. Pr\u00e9hist. Fr. 9, 700\u2013709 (1912).<\/p>\n Martin, H. L.\u2019Homme Fossile de la Quina (Libraire Octave Doin, 1923).<\/p>\n Pap, I., Tillier, A. M., Arensburg, B. & Chech, M. The Subalyuk Neanderthal remains (Hungary): a re-examination. Ann. Hist. Nat. Mus. Natl Hung. 88, 233\u2013270 (1996).<\/p>\n Sanchez, F. Comparative biometrical study of the Mousterian mandible from Cueva del Boquete de Zafarraya (M\u00e1laga, Spain). J. Hum. Evol. 14, 125\u2013138 (1999).<\/p>\n Article<\/a>\u00a0 Vl\u010dek, E. Fossile Menschenfunde von Weimar-Ehringsdorf, Weimarer Monographien zur Ur- und Fr\u00fchgeschichte Vol. 30 (Landesamt f\u00fcr Arch\u00e4ologische Denkmalpflege, 1993).<\/p>\n Condemi, S. Les N\u00e9andertaliens de La Chaise: Abri Bourgeois-Delaunay (CTHS, 2001).<\/p>\n Bar-Yosef, O. & Vandermeersch, B. (eds) Le Squelette Moust\u00e9rien de K\u00e9bara 2 (CNRS, 1991).<\/p>\n Arambourg, C. & Hoffstetter, R. Le Gisement de Ternifine Vol. 1 (Masson, 1963).<\/p>\n Hooton, E. A., Hencken, H. O. & Snow, C.hE. The ancient Palestinian: Skh\u016bl V reconstruction. Am. School Prehist. Res. 17, 5\u201310 (1953).<\/p>\n Sollas, W. J. The Chancelade skull. J. R. Anthropol. Inst. 57, 89\u2013122 (1927).<\/p>\n Martin, H. Caract\u00e8res des squelettes humains quaternaires de la vall\u00e9e du Roc (Charente). Bull. Mem. Soc. Anthropol. Paris 8, 103\u2013129 (1927).<\/p>\n Article<\/a>\u00a0 Vercellotti, G., Alciati, G., Richards, M. P. & Formicola, V. The Late Upper Paleolithic skeleton Villabruna 1 (Italy): a source of data on biology and behavior of a 14,000year-old hunter. J. Anthropol. Sci. 86, 143\u2013163 (2008).<\/p>\n PubMed<\/a>\u00a0 Formicola, V. Una mandibola umana dal deposito dell\u2019Epigravettiano finale delle Arene Candite (scavi 1970). Rev. Antropol. 64, 271\u2013278 (1986).<\/p>\n Odano, A. M. & Riquet, R. Le gisement pr\u00e9historique de Dar-es-Soltane 2. Champ de tir de El Menzeh \u00e0 Rabat (Maroc). Note pr\u00e9liminaire. 2-\u00c9tude anthropologique des restes post-at\u00e9riens. Bull. d\u2019Arch\u00e9ologie Maroc. 11, 25\u201363 (1978).<\/p>\n Deb\u00e9nath, A. Nouveaux restes humains at\u00e9riens du Maroc. C. R. Acad. Sci., Paris 290, 851\u2013852 (1980).<\/p>\n MathSciNet<\/a>\u00a0 Crognier, E. & Dupouy-Madre, M. Les Natoufiens du Nahal Oren (Ouadi Fallah). Etude anthropologique. Pal\u00e9orient 2, 103\u2013121 (1974).<\/p>\n Article<\/a>\u00a0 Henke, W. Vergleichend-morphologische Kennzeichnung der Jungpal\u00e4olithiker von Oberkassel bei Bonn. Z. Morphol. Anthropol. 75, 27\u201344 (1984).<\/p>\n Article<\/a>\u00a0 Hershkovitz, I. et al. Ohalo II H2: a 19,000-year-old skeleton from a water-logged site at the Sea of Galilee, Israel. Am. J. Phys. Anthropol. 96, 215\u2013234 (1995).<\/p>\n Article<\/a>\u00a0 Soficaru, A., Dobos, A. & Trinkaus, E. Early modern humans from the Pestera Muierii, Baia de Fier, Romania. Proc. Natl Acad. Sci. USA 103, 17196\u201317201 (2006).<\/p>\n Article<\/a>\u00a0 Crevecoeur, I. Etude Anthropologique des Restes Humains de Nazlet Khater (Pal\u00e9olithique Superieur, Egypte). PhD Thesis, Universit\u00e9 Sciences et Technologies Bordeaux I (2006).<\/p>\n Thoma, A. Morphology and affinities of the Nazlet Khater man. J. Hum. Evol. 13, 287\u2013296 (1984).<\/p>\n Article<\/a>\u00a0 Anderson, J. E. in The Prehistory of Nubia Vol. 2 (ed. Wendorf, F.) 996\u20131040 (Southern Methodist Univ. Press, 1968).<\/p>\n Crevecoeur, I. From the Nile to the Danube: a comparison of the Nazlet Khater 2 and Oase 1 early modern human mandibles. Anthropologie 42, 203\u2013213 (2004).<\/p>\n Trinkaus, E. et al. An early modern human from the Pestera cu Oase, Romania. Proc. Natl Acad. Sci. USA 100, 11231\u201311236 (2003).<\/p>\n Article<\/a>\u00a0 Trinkaus, E. & Svoboda, J. (eds) Early Modern Human Evolution in Central Europe. The People of Doln\u00ed V\u011bstonice and Pavlov. The Dolni Vestonice Studies Vol. 12 (Oxford Univ. Press, 2006).<\/p>\n Sl\u00e1dek, V., Trinkaus, E., Hillson, S. W. & Holliday, T. W. The People of the Pavlovian. Skeletal Catalogue and Osteometrics of the Gravettian Fossil Hominids from Doln\u00ed V\u011bstonice and Pavlov. The Dolni Vestonice Studies Vol. 5 1\u2013244 (Brno, 2000).<\/p>\n Drozdov\u00e1, E. The evaluation of a rediscovered fragment of human lower jaw, No 21 from P\u0159edmost\u00ed u P\u0159erova. Archeol. Rozhl. 53, 452\u2013460 (2001).<\/p>\n Dutour, O. Palimpseste pal\u00e9oanthropologique sur l\u2019\u2018Homme fossile d\u2019Asselar\u2019 (Sahara). Pr\u00e9hist. Anthropol. M\u00e9dit. 1, 73\u201383 (1992).<\/p>\n Gambier, D. Vestiges humains du gisement d\u2019Isturitz (Pyr\u00e9n\u00e9es-Atlantiques): \u00e9tude anthropologique et analyse des traces d\u2019action humaine intentionnelle. Antiqu. Natl. 22\u201323, 9\u201326 (1990).<\/p>\n Wood, B. Koobi Fora Research Project, Vol. 4: Hominid Crania Remains (Clarendon, 1991).<\/p>\n Rightmire, G. P. The Evolution of Homo Erectus: Comparative Anatomical Studies of an Extinct Human Species (Cambridge Univ. Press, 1990).<\/p>\n Wollny, G. et al. MIA\u2014a free and open-source software for gray scale medical image analysis. Source Code Biol. Med. 8, 20 (2013).<\/p>\n Article<\/a>\u00a0 Cannoodt, R. & Bengtsson, H. rcannood\/princurve: princurve 2.1.4. Zenodo https:\/\/doi.org\/10.5281\/zenodo.3351282<\/a> (2019).<\/p>\n Davies, T. W. tomdavies15\/EDJ_GM: geometric morphometric analysis of enameldentine junction morphology. Zenodo https:\/\/doi.org\/10.5281\/zenodo.10255289<\/a> (2023).<\/p>\n Dryden, I. & Mardia, K. V. Statistical Shape Analysis (Wiley, 1998).<\/p>\n Molnar, S. Human tooth wear, tooth function and cultural variability. Am. J. Phys. Anthropol. 34, 175\u2013189 (1971).<\/p>\n Article<\/a>\u00a0 Turner, C., Nichol, C. & Scott, G. in Advances in Dental Anthropology (eds Kelley, M. & Larsen, C.) 13\u201331 (Wiley Liss, 1991).<\/p>\n Skinner, M. M. et al. Dental trait expression at the enamel-dentine junction of lower molars in extant and fossil hominoids. J. Hum. Evol. 54, 173\u2013186 (2008).<\/p>\n Article<\/a>\u00a0 Bailey, S., Skinner, M. M. & Hublin, J. J. What lies beneath? An evaluation of lower molar trigonid crest patterns based on both dentine and enamel expression. Am. J. Phys. Anthropol. 145, 505\u2013518 (2011).<\/p>\n Article<\/a>\u00a0 Bailey, S. E. Neandertal Dental Morphology: Implications for Modern Human Origins. PhD thesis, Arizona State University (2002).<\/p>\n Burnett, S. E., Hawkey, D. E. & Turner, C. G. II Population variation in human maxillary premolar accessory ridges (MxPAR). Am. J. Phys. Anthropol. 141, 319\u2013324 (2010).<\/p>\n Article<\/a>\u00a0 Benazzi, S. et al. Early dispersal of modern humans in Europe and implications for Neanderthal behaviour. Nature 479, 525\u2013528 (2011).<\/p>\n Article<\/a>\u00a0 Benazzi, S. et al. Comparison of dental measurement systems for taxonomic assignment of Neanderthal and modern human lower second deciduous molars. J. Hum. Evol. 61, 320\u2013326 (2011b).<\/p>\n Article<\/a>\u00a0 Benazzi, S. et al. Cervical and crown outline analysis of worn Neanderthal and modern human lower second deciduous molars. Am. J. Phys. Anthropol. 149, 537\u2013546 (2012).<\/p>\n Article<\/a>\u00a0 Bailey, S. E., Benazzi, S., Buti, L. & Hublin, J. J. Allometry, merism, and tooth shape of the lower second deciduous molar and first permanent molar. Am. J. Phys. Anthropol. 159, 93\u2013105 (2016).<\/p>\n Article<\/a>\u00a0 Bailey, S. E., Sorrentino, R., Mancuso, G., Hublin, J. J. & Benazzi, S. Taxonomic differences in deciduous lower first molar crown outlines of Homo sapiens and Homo neanderthalensis. J. Hum. Evol. 147, 102864 (2020).<\/p>\n Article<\/a>\u00a0 Jolliffe, I. T. Principal Component Analysis https:\/\/doi.org\/10.1007\/b98835<\/a> (Springer, 2002).<\/p>\n Le Cabec, A., Gunz, P., Kupczik, K., Braga, J. & Hublin, J.-J. Anterior tooth root morphology and size in Neanderthals: taxonomic and functional implications. J. Hum. Evol. 64, 169\u2013193 (2013).<\/p>\n Article<\/a>\u00a0 Meyer, M. R. & Haeusler, M. Spinal cord evolution in early Homo. J. Hum. Evol. 88, 43\u201353 (2015).<\/p>\n Article<\/a>\u00a0
\n ADS<\/a>\u00a0
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\n CAS<\/a>\u00a0
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\n PubMed<\/a>\u00a0
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\n PubMed<\/a>\u00a0
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\n ADS<\/a>\u00a0
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\n PubMed<\/a>\u00a0
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\n PubMed<\/a>\u00a0
\n
\n Google Scholar<\/a>\u00a0\n <\/p>\n
\n PubMed<\/a>\u00a0
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\n Google Scholar<\/a>\u00a0\n <\/p>\n
\n PubMed<\/a>\u00a0
\n CAS<\/a>\u00a0
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\n PubMed<\/a>\u00a0
\n
\n Google Scholar<\/a>\u00a0\n <\/p>\n
\n PubMed<\/a>\u00a0
\n
\n Google Scholar<\/a>\u00a0\n <\/p>\n","protected":false},"excerpt":{"rendered":"Meyer, M. et al. Nuclear DNA sequences from the Middle Pleistocene Sima de los Huesos hominins. Nature 531,…\n","protected":false},"author":2,"featured_media":394079,"comment_status":"","ping_status":"","sticky":false,"template":"","format":"standard","meta":{"footnotes":""},"categories":[32],"tags":[9739,38767,1159,1160,79],"class_list":["post-394078","post","type-post","status-publish","format-standard","has-post-thumbnail","category-science","tag-anthropology","tag-geomagnetism","tag-humanities-and-social-sciences","tag-multidisciplinary","tag-science"],"_links":{"self":[{"href":"https:\/\/www.newsbeep.com\/us\/wp-json\/wp\/v2\/posts\/394078","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/www.newsbeep.com\/us\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/www.newsbeep.com\/us\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/www.newsbeep.com\/us\/wp-json\/wp\/v2\/users\/2"}],"replies":[{"embeddable":true,"href":"https:\/\/www.newsbeep.com\/us\/wp-json\/wp\/v2\/comments?post=394078"}],"version-history":[{"count":0,"href":"https:\/\/www.newsbeep.com\/us\/wp-json\/wp\/v2\/posts\/394078\/revisions"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/www.newsbeep.com\/us\/wp-json\/wp\/v2\/media\/394079"}],"wp:attachment":[{"href":"https:\/\/www.newsbeep.com\/us\/wp-json\/wp\/v2\/media?parent=394078"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/www.newsbeep.com\/us\/wp-json\/wp\/v2\/categories?post=394078"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/www.newsbeep.com\/us\/wp-json\/wp\/v2\/tags?post=394078"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}