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Sophia Laukli (far left) has made her mark in the endurance sports world. Training for the events in which she specializes has become a mixture of science, intuition, and discipline. (Photo: NordicFocus)
Over the past few years, I’ve been studying the work of Herman Pontzer, PhD, whose research in human energetics challenges long-held paradigms of how endurance athletes adapt to training. For decades, endurance training theory was based on the principle of cumulative energy expenditure: train more, eat more, adapt more, and perform better. Pontzer’s Constrained Total Energy Expenditure (TEE) model calls this theory into question.
His concept suggests: the body does not endlessly increase energy production with ever increasing physical activity. Instead, total daily energy expenditure adapts by reallocating energy away from other physiological systems. For endurance athletes, this represents a paradigm shift with direct implications for health, optimal training load, recovery, and long-term adaptation.
While we await further research, the compelling research from Pontzer should give us pause to reconsider how we approach training athletes at all levels.
The Constrained Energy Model: Key Components
Pontzer breaks down energy use into three primary categories:
Basal Metabolic Rate (BMR): energy required for basic functions such as cellular repair, recovery, and adaptation.
Physical Activity Energy Expenditure (PAEE): the energy cost of exercise.
Thermic Effect of Food (TEF): energy required for digestion and metabolism.
Pontzer expresses Constrained Total Energy Expenditure (TEE) as multiples of BMR to make meaningful comparisons across individuals and populations. For a point of reference, in my experience most endurance athletes should target BMR multipliers in the 1.7-2.0 TEE multiplier range
Population/ Activity Level
TEE Multiplier (TEE ÷ BMR)
Notes
Sedentary
1.4–1.6
Desk job, little structured activity
Moderately active
1.7–2.0
Regular exercise, recreational athletes
Hunter-gatherers (Hadza)
2.0
High activity, but within sustainable range
Endurance athletes (highly trained)
2.2–2.5+ (Temporarily)
Heavy training blocks or racing periods
Tour de France ride
3.5–4.5 (short-term)
Only sustainable for weeks, not year-round
Constrained ceiling (long-term sustainable limit)
~2.5
Beyond this, the body compensates by reducing other functions and create health consequences. (immune, reproductive, endocrine)
Even extremely active groups like the Hadza tribe who Pontzer initially studied, or ultra-endurance athletes rarely exceed ~2.5× BMR over the long term. Short-term spikes above this ceiling are possible, but the body compensates by down-regulating other functions—immune, hormonal, and reproductive—to stay within limits.
For example, Tour de France riders may reach 3.5–4.5× BMR during the race, but their long-term average settles back into the 2–2.5× range. After the Tour, riders require extended recovery to normalize immune and endocrine function, evidence of the physiological “borrowing” predicted by the constrained model.
Why It Matters for Endurance Athletes
Limits to “More is Better”
The TEE model suggests that beyond a certain point, piling on more training volume does not yield proportionally greater adaptation. Excessive energy demand forces trade-offs: suppressed endocrine and immune function, impaired recovery, stagnation, or maladaptation.
Athletes may look “fit but fragile”: capable of high short-term performance but prone to illness, overtraining, or injury in the long run.
A specific example
A 16-year-old girl, 5’5’’ and 130 lb has a BMR of ~1380 kcal/day. Assuming a high level of activity and a 2.2 TEE multiplier, the TEE is about 3036 kcals per day. Subtracting the BMR leaves a maximum of 1650 Kcals per day for training and other energy expenditures.
Using a conservative estimate of all training energy expenditure the available energy amounts to about 11-12-hours per week of endurance training and perhaps an hour less if there is a high intensity component. That is the very highest energy expenditure this female athlete can maintain, adapt to training, and remain healthy. Any training load above has to be repaid in the short term and is not sustainable for other than brief periods,from a health, adaptation, or performance perspective. To be clear, training more than a dozen hours per week is probably not optimal for this athlete. From what I have seen over the last 20 years, the HRV data supports this conclusion.
For endurance athletes
For reference in our assessments using Firstbeat Technology, low intensity training, Level 1 will require 8-10 Kcals per minute, Moderate Intensity Training Level 3, 12-14 kcals per minute, and High Intensity Training Level 4, 15-18 kcals per minute. Of course, individual expenditures will vary depending on the athlete’s efficiency and fitness.
Training consistently beyond at TEE of 2.5 BMR, often means hidden physiological costs: menstrual dysfunction, chronic inflammation, suppressed immunity, and poor recovery.
The TEE model provides the mechanistic ceiling: humans rarely sustain >2.5× BMR without consequences.
RED-S documents the clinical outcomes: the health and performance impairments seen when athletes operate beyond this ceiling.
For endurance athletes and coaches, the overlap is clear, respecting the constrained energy ceiling is central to preventing RED-S. Sustainable performance requires balancing workload with recovery, nutrition, and life stress to avoid the silent drift into energy deficiency.
Recovery as a Competitive Advantage
If high training loads divert energy from critical systems, then recovery is not optional, it’s essential. Sleep, nutrition, low-intensity activity, and genuine rest are the practices that restore compromised systems and sustain adaptation.
In this light, moderate but consistent training paired with high-quality recovery may prove more effective long term than chronic high-volume training with poor recovery.
Individual Variability
The constrained ceiling is not the same for everyone. Age, sex, metabolic rate, training history, and life stressors all shift the point where trade-offs occur.
Young, hormonally robust male athletes may tolerate higher loads.
Female athletes with low energy availability may reach the ceiling much sooner.
This underscores the importance of individualized monitoring tools, HRV, menstrual cycle tracking, or biomarker testing, to gauge stress and recovery.
Implications for Periodization
Pontzer’s work provides a biological basis for adaptive periodization: alternating blocks of high training stress with recovery phases. This aligns with polarized or pyramidal models already used successfully by many elite endurance athletes.
Also Important, “life stress” also counts toward the energy budget. Academic, work, or family stress draws from the same pool as training, reinforcing the need to coach the whole athlete, not just their training log.
Practical Application
How coaches and athletes can integrate Pontzer’s framework:
Estimate BMR – use predictive equations like Mifflin–St Jeor, or ideally indirect calorimetry.
Track TEE – wearable tech (Garmin, WHOOP, Firstbeat) offers estimates; research uses doubly-labeled water.
Calculate TEE/BMR – monitor when ratios exceed ~2.3–2.5 for multiple days.
Adjust Training – treat sustained values above the ceiling as a warning sign of overtraining risk and an opportunity to prioritize recovery.
Conclusion: Smarter Training Within Limits
The constrained energy model forces us to rethink the long-held belief that more training always equals more adaptation. The body operates within a finite energy budget. Of course, we have not covered the importance of fueling and nutrition which is emerging as well. Sustainable performance depends not only on the training load and stimulus but also on respecting the body’s ability to remain healthy and in energy balance.
Pontzer’s research validates what many experienced coaches and athletes have long understood, the science and art of endurance training lies not in endlessly pushing, but in balancing stimulus with recovery. In the space between effort and adaptation lies the true craft, and science, of coaching.